Max Krummel received his undergraduate degree in biology and chemistry at the Univeristy of Illinois Urbana-Champaign. He received his Ph.D. at UC Berkeley working in the laboratory of James Allison, where he was involved in early experiments demonstrating that the T cell protein CTLA-4 acts as a checkpoint inhibitor, work that helped lay the foundation for the most important class of modern cancer immunotherapy drugs. After a postdoctoral fellowship with Allison, he completed a second postdoc at the Walter and Eliza Hall Institute in Melbourne, Australia, working with Bill Heath and Ken Shortman on dendritic cell biology. He then completed a third postdoctoral fellowship with Mark Davis at Stanford University before joining UCSF in 2001. He is faculty director of UCSF’s Biological Imaging Development Center.