Ophir Klein received his undergraduate degree from UC Berkeley, and M.D. and Ph.D. degrees from Yale School of Medicine. After completing his residency in pediatrics at Yale, Klein completed a second residency, in genetics, at University of California, San Francisco. In 2014 he was named the Larry L. Hillblom Distinguished Professor in Craniofacial Anomalies, and in 2016, the Charles J. Epstein Professor of Human Genetics at UCSF. He serves as chief of UCSF’s Division of Medical Genetics, chair of the Division of Craniofacial Anomalies, medical director of the Craniofacial Center, and director of the Program in Craniofacial Biology.